Primary-Secondary Growth-Stems-2

HoyaXsec.jpg (86724 bytes)
Lecture Directory

Coleus-3.jpg (75322 bytes)
Prior Page

woodring.jpg (51247 bytes)
Next Page

Secondary Growth

Secondary growth is due to the action of Lateral Meristems These include the Vascular Cambium in Shoots and Roots of Gymnosperms and Dicots, the Secondary Thickening Meristem in Monocots and the Cork Cambium which occurs in all of the above.

Secondary growth increases the girth of plant organs, This is most obvious in the shoots of large trees which are wide at the base and taper towards the apex. Few plants would attain a height of a few meters without secondary growth in their stems.

Secondary growth is less obvious in roots. However, cursory observations of plants on campus or in nature will quickly verify the presence of secondary growth in the form of Periderm (cork). The thickness of these roots can only be due to the formation of secondary vascular tissues.  Roots play an enormous role in supporting shoot systems. Look at the bases of the Giant Sequoia trees below. Their stems are enlarged at the base but without an enormous root system the stems would topple.

We will concentrate on stems now, and will examine roots later.

SequoiaBaseCrop.jpg (32544 bytes)
Base of Giant Sequoia Trees - Note the enlarged basal areas of the stem. One can only imagine the size and complexity of the underlying root system which is largely due to secondary growth.

woodrings.jpg (56469 bytes)
Wood is produced by the Vascular Cambium. Wood is one of the most useful natural products on the planet. It is still used as fuel in most parts of the world. The trees which produce it have enormous importance due to their effect on local and global climates.

PeriScaleBark240.jpg (50367 bytes)

PeridermScaleBark240.jpg (34717 bytes)

Internal secondary growth increases the diameter of stems and roots. The Cork Cambium produces  Periderm which covers the surface of expanding stems and roots.  Periderm prevents water loss and protects the pant from environmental and pathological insults.

rootclif260.jpg (19310 bytes)
Secondary growth in roots is usually obscured by the soil. However, the roots of this plant are displayed against the cliff face above. Careful scrutiny would show that these roots had secondary growth in the form of periderm and vascular tissues.

LatRootPandanus300.jpg (27710 bytes)
Lateral root of hala (Pandanus) The surface layer is Periderm. The protrusions are lateral root primordia. Pandanus is a monocot.

Page Top