A FORMAL CLASSIFICATION OF THE CYCADS
The classification of the extant cycads proposed by Dennis Stevenson in
1990 (Memoirs of the New York Botanical Garden 57:8-55) has been formalised
by Dennis in a publication dedicated to the memory of Sergio Sabato (Brittonia 44(2),
in press). The classification places the 11 genera and 3 families in a hierarchical
structure based on cladistic analyses of morphological, anatomical, karyological,
physiological and phytochemical data, and is summarised by the following diagram:
Order: Cycadales
Suborder: Cycadineae
Family: Cycadaceae
Genus: Cycas [Type: C. circinalis L.]
Suborder: Zamiineae
Family: Stangeriaceae
Subfamily: Stangerioideae
Genus: Stangeria [Type: S. eriopus (Kunze) Baillon]
Subfamily: BowenioideaeGenus: Bowenia [Type: B. spectabilis Hook. ex Hook. f.]
Family: ZamiaceaeSubfamily: Encephalartoideae
Tribe: Diooeae
Genus: Dioon [Type: D. edule Lindley]
Tribe: EncephalarteaeSubtribe: Encephalartinae
Genus: Encephalartos [Lectotype: E. friderici-guilielmi Lehmann]
Subtribe: Macrozamiinae
Genus: Macrozamia [Lectotype: M. riedlei(Fishcer ex
Gaudichaud) C.A. Gardner]
Genus: Lepidozamia [Type: L. peroffskyana Regel]
Subfamily: Zamioideae
Tribe: Ceratozamieae
Genus: Ceratozamia [Type: C. mexicana Brongn.]
Tribe: ZamieaeSubtribe: Microcycadinae
Genus: Microcycas [Type: M. calocoma (Miquel) A. DC.]
Subtribe: Zamiinae
Genus: Zamia [Type: Z. pumila. L.]
Genus: Chigua [Type: C. restrepoi E. Stevenson]
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