Monocot Roots

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CornWholeStele240.jpg (39872 bytes)

CornRootStele240.jpg (15120 bytes)

Vascular Tissues in a Corn Root - The cells with the largest Diameters are Vessel Members. Interface of the Stele & the Cortex-Follow the Cortical Parenchyma from the top of the image to the first cells which have thick, red-stained walls. These form  the Endodermis.

The Endodermis usually develops exceedingly thick secondary walls at levels in the root which are no longer absorbing water. This obscures the Casparian Strips but makes the Endodermis more obvious.

The organization of Monocot roots like Sugarcane is similar to that found in dicots like Ranunculus. However, tracheary elements may be absent from the center of the root. There are usually more xylem arms in monocots. The number of arms is indicated by the following terms.

Number of Xylem Arms  

Two = Diarch
Three = Triarch
Four = Tetrarch
Five = Pentarch
Six   = Hexarch
More than five = Polyarch

Monocots are usually Polyarch. Bundles of Phloem alternate with the Xylem arms. The xylem contains many lignified cells in older portions of the root. The phloem stands out because its cells have thin, unlignified cell walls.

ClintoniaRootX-S240.jpg (74675 bytes)

Cross-section of Clintonia (monocot) Root. Count the number of xylem arms.

RotSmilaxWholXS300.jpg (40873 bytes)

RotSmilaxEpi-Xylm300.jpg (19191 bytes)

Cross-section of Smilax (monocot) Root. The tracheary elements of the Xylem are the cells with the largest diameters. Count the number of xylem arms.
The Endodermis is the densely stained, lignified layer which surrounds the Stele.

SmilaxRotMaturePhloemXylem300.jpg (41929 bytes)
Smilax Stele showing bundles of Phloem  alternating with Xylem Vessel Members.

The cell pattern is distorted by the extreme enlargement of the  Vessel Members.

Smilax root at higher magnification. Locate the Xylem and Phloem as well as the Endodermis, Cortex and Epidermis.

 

 

 

 

SmilaX-sEpiCortCasp300.jpg (51266 bytes)
Smilax root showing the Stele, Endodermis, Cortex, Exodermis & Epidermis. The Exodermis has the same anatomy as the Endodermis and probably restricts water loss to the outside in older roots.

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