XVI International Botanical Congess
The diplolepideous-alternate peristome at its most highly developed has endostome segments attached to a basal membrane and positioned alternate to the outer exostome teeth, with cilia between the segments and opposite to the exostome teeth. This peristome type defines the Bryidae, which includes four orders, namely the Bryales, Leucodontales, Hypnales and Hookeriales, of which the latter three are exclusively pleurocarpous in their growth form. Analyses of chloroplast (rbcL, rps4 and trnL) and nuclear (18S rRNA) gene sequences strongly support the paraphyly of the Bryidae, with both the Splachnidae and Orthotrichidae having possible independent origins from within the Bryidae. The three pleurocarpous orders also form a monophyletic group.